A new species of Oeneis from Alaska, United States, with notes on the Oeneis
Andrew D. Warren(1), Shinichi Nakahara(1), Vladimir A. Lukhtanov(2,3), Kathryn M. Daly(4), Clifford D.
Ferris (5), Nick V. Grishin(6), Martin Cesanek(7) and Jonathan P. Pelham(8)
The Journal of Research on the Lepidoptera
ISSN 0022-4324 (print)
Abstract. Oeneis tanana A. Warren & Nakahara is described from the Tanana River Basin in
southeastern Alaska, USA. This new taxon belongs to the bore group of Oeneis Hübner,  and is
apparently closest to O. chryxus (E. Doubleday, ) by morphology, including its larger size and
similarity of the female genitalia. In wing patterns and COI mitochondrial DNA barcode sequences,
it is reminiscent of O. bore (Esper, 1789). A review of O. chryxus subspecies suggest that some may
Key words: Beringia, butterflies, cryptic species, hybrid species, Nearctic, speciation, taxonomy, Yukon Territory.
Received: 21 December 2015
Published online at www.lepidopteraresearchfoundation.org on 15 March 2016